Browsing by Author "Zacharias, Peter John Kenneth."

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Aspects of the invasion of southern tall grassveld by Aristida junciformis subsp. junciformis Trin. et Rupr.
(1998) Van Zyl, Douglas Dirk.; Granger, James Edmund.; Zacharias, Peter John Kenneth.
Aristida junciformis subsp. junciformis Trin. et Rupr. is an indigenous, densely tufted, evergreen perennial grass associated with the degeneration of grasslands over large areas of South Africa. More than two hundred thousand square kilometres of veld, c. 17% of the total land area of South Africa, contains A. junciformis. The aim of this study was to improve our understanding of the mechanisms by which this species invades and dominates grassland, specifically in this study, Southern Tall Grassveld. Aristida junciformis also has a low nutritional value resulting in a very low grazing value. The unpalatability of the plant is due to the high tensile strength of the leaves, whose hard, fibrous laminas are very difficult for a grazing animal to crop once their length attains 30 cm or more. This species lack of response to conventional grazing practices has often resulted in a grassland in which the carrying capacity has diminished to such an extent as to be virtually useless for grazing in a relatively short time. The extent of encroachment of veld by A. junciformis appears dependant on the frequency of disturbance and rest afforded to the veld. Annual burning and mowing maintained the species composition of A. junciformis at levels <10% whilst protection from fire, burning or grazing allows this species to dominate the herbaceous layer at levels approaching 90%. Aristida junciformis does not become moribund and is unlikely to die if left undefoliated. As the abundance of A. junciformis increases, veld deterioration gradually accelerates through increased selective grazing on remaining palatable species. Competition from adult A. junciformis plants increases the mortality of seedlings and constrains growth of surviving seedlings. Tiller production of Aristida junciformis seedlings declined from an average of 5.2 tillers per seedling in a no competition situation to 2.2 tillers per seedling when subjected to full competition. Tiller production of T. triandra seedlings decreased from 9.6 tillers per seedling free from competition to 3.3 tillers per seedling subject to full competition for resources. Once these seedlings have become adult plants they are avoided by grazing animals and grazing pressure on the remaining palatable species consequently increases. This allows the A. junciformis plants to increase their size and density in the sward. Once this density is sufficiently high, grass seedlings of either A. junciformis or more desirable grass species such as T. triandra are unable to establish, eventually resulting in a monospecific stand of A. junciformis if left undisturbed. Large amounts of caryopses are produced by A. junciformis - up to a 19 000 caryopses from a large mature plant (c. 38 000 caryopses/m²). Of these c. 40% is likely to be infertile but the remaining c. 60% viable caryopses are dispersed in a typically leptokurtic distribution, the number of caryopses dispersed rapidly declining within a 10 metre radius. A large proportion of the caryopses was trapped in surrounding foliage but in open swards caryopses had greater opportunity to be blown further distances than in a closed sward. The density of A. junciformis caryopses on the soil surface was positively correlated with the density of flowering adult plants in the area and varied from 400 caryopses/m² (density of parent plants c. 0.6 plants/m²) in less effected areas to 11000 caryopses/m² (density of adult plants c. 6 plants/m²) in severely encroached areas. The primary function of the three awns appears to be orientating the caryopsis correctly for in its descent from the parent plant to expedite germination. Caryopses orientated vertically with the awns uppermost exhibited the highest and most rapid germination (67%) compared to caryopses lying horizontally (35%) whilst only 1% of inverted caryopses germinated. Caryopses trapped in litter and effectively held off the soil surface failed to germinate. Removal of the glumes from A junciformis seed greatly enhanced the rate and overall germination of the seed except for inverted seed of which <1% germinated. The highest numbers of A. junciformis seedlings (32 seedlings/m²) were found in those areas with the highest density of caryopses on the soil surface. Despite the large amounts of caryopses produced, dispersed and landing in apparently suitable micro-sites for germination, comparatively few A. junciformis seedlings (n=992) were found and overall germination ranged between 1 % and 4% of the initial caryopses density on the soil surface. Seedling survival through winter was low with only 13% surviving to the following spring. The basal areas of A. junciformis increased overall by 66% whilst that of other grass species increased overall by only 3% in the time monitored. These results suggest that the primary method of encroachment of A. junciformis in the grass sward appears to be through vegetative expansion and not seedling recruitment. Frequent defoliation of the sward and avoidance of overgrazing to enhance the competitive abilities of palatable species and provide as high a fuel load as possible appear to be the most economically and logistically feasible ways to remove or at least inhibit A. junciformis veld encroachment at present.
A comparative classification of the sourish-mixed bushveld on the farm Roodeplaat (293 JR) using quadrat and point methods.
(1995) Panagos, Michael David.; Westfall, Ribert Howard.; Zacharias, Peter John Kenneth.; Ellis, Roger.
An area and a point-based technique were used together at each of the same 75 sampling sites (stands), on a Sourish-Mixed Bushveld farm, to collect data for the classification and mapping of the vegetation. Both sets of data were synthesized using the same computer program package and the efficacy of the resulting classifications as well as the efficiency of the two field sampling techniques was compared. Following this, a continuous 7 752 point (1 m apart) transect was carried out, traversing the farm, in order to determine the optimum scales at which to sample Sourish-Mixed Bushveld so as to increase classification efficacy and improve community boundary recognition. The results indicated that (1) the arbitrarily chosen sampling scale of 1:8 000 was too large for "farm-scale" studies; (2) the area-based method proved to be satisfactory in that the classification and vegetation map produced with this method were verified spatially and environmentally; (3) the point-based method was deficient as a classificatory and mapping tool at large scales, since too few species were recorded with this method to make any sense of the classification and mapping of the vegetation was not possible; (4) less time per species was spent using the area-based method but because more species per stand were recorded with this method, the point-based method was quicker per stand; (5) the area-based method was easier to use in dense vegetation and irregular terrain; and (6) the optimum sampling scales for Sourish-Mixed Bushveld, as indicated by the synthesis of the continuous transect data, are about 1:12 000, 1:50 000 and 1:250 000.
A comparison of in-field techniques for estimating the feed intake of young boer goats on a Leucaena leucocephala/grass hay diet.
(1997) Letty, Brigid Aileen.; Zacharias, Peter John Kenneth.; Morris, Craig Duncan.
Two methods of estimating the intake of a 25% leucaena : 75% grass hay diet by young male Boer goats were assessed, (a) The purine derivative technique which uses the urinary excretion of purine derivatives (expressed relative to creatinine concentration in the same sample) as an index of feed intake, and (b) the conventional marker method, utilizing chromic oxide (Cr₂O₃) contained in gelatin capsules and dosed twice daily, as the marker. Following a prerun the two techniques were compared in three runs of an indoor experiment. In each run 10 goats were randomly allocated to five feeding levels (500 to 1100 g fodder d ¯¹ on air dried basis). A preliminary and an adaptation period during which goats were dosed with the Cr₂O₃ and fed their daily feed allowance, was followed by a 4 day collection period during which spot samples of urine were collected and analysed for allantoin and creatinine (allantoin being used instead of total PDs) and faecal samples were collected for chromium analysis and percentage dry matter determination. For the first two runs, two grab samples per day for each goat were bulked and analysed for chromium content. For the last run, the total daily faecal collection was subsampled and analysed for chromium. Work was done in metabolic crates to determine the effect of time of collection on the ratio of allantoin : creatinine (A/C) in spot urine samples and it was found to non-significant (P>0.05). Linear regressions of: (a) feed intake expressed per unit metabolic mass (g.d ¯¹.LW ¯°∙⁷⁵ (I_mmass)) against A/C ratio; (b) faecal output (g d ¯¹) against feed intake (g d ¯¹); and (c) faecal chromium concentration (mg kg ¯¹) against faecal output (g d ¯¹) were fitted to the data. During the prerun, only regression (a) was fitted and was non-significant (P>0.05), showing no trend at all. For the first true run, the regression of I_mmass against A/C ratio was significant and the correlation was high (P≤ 0.001, R² [A] 0.715, n = 10) but for the second and third runs, the correlations only became significant when the apparent outliers were discarded from the data. (Run 2: P≤ 0.001, R² [A] 0.824, n = 8; Run 3: P≤ 0.05, R² [A] 0.430, n = 9). It was concluded that the relation between I_mmass and A/C ratio is not well enough defined to be used for predictive purposes. When regression (b) was investigated, all the runs produced significant results (P≤0.001, P≤0.01, P≤ 0.001 for runs 1,2 and 3 respectively) however the correlations were not as high as expected (R² [A] being 0.714, 0.565 and 0.863 respectively). For the regression of faecal Cr concentration against faecal output (regression c), all runs showed significant relations (P< 0.001, P< 0.0001, P:s 0.001 for runs 1,2 and 3 respectively) and the correlations were high (R²[A] being 0.836, 0.837 and 0.912 respectively). The data from the three runs were pooled and single equations established for regressions (b) and (c) to allow for the prediction of intake from faecal chromium concentration. Faecal output = feed intake * 0.448 + 19.341 (P≤ 0.001, r 0.853, R² [A] 0.718, SE 25.664, n - 30) Faecal chromium concentration = faecal output * -241.547 + 1.315E+05 (Ps 0.001, r 0.904, R² [A] 0.811, SE 5603.788, n = 30). In vitro figures were determined for a range of leucaena : hay mixes but no apparent trend was found between percentage leucaena in the mix and the digestibility of the mix. These results compared favourably with in vivo results obtained for a 25% leucaena : 75% hay mix. Neither technique proved entirely satisfactory, but the external marker method was found to be more effective than the purine derivative technique. More work is required especially with respect to the latter method.
Determinants of grass production and composition in the Kruger National Park.
(2003) Zambatis, Nicholas.; Zacharias, Peter John Kenneth.; Morris, Craig Duncan.; Biggs, Harry Cawood.
The dynamics and complexities of climate-soil-vegetation relations in the Kruger National Park are poorly known. Although primary production and composition of the grass layer are very important components of the Park's ecosystem, equally little is known about the determinants of these parameters. A better understanding of these processes and relations will be of value to the management of this Park, as well as providing a better insight into these complex dynamics. A study was consequently undertaken covering a 14-year period to identify the most important determinants of above-ground grass production and composition. At the core of the study is the soil water balance. The use of evapotranspiration data in a study of this nature is however not absolutely essential, provided a variety of rainfall parameters are used, though it has the important advantage of providing a much more detailed and more complete insight into the relations of the grass sward with its environment. Stepwise and tree regression procedures were used to identify the important factors. It is concluded that rainfall in its various forms is the primary determinant of grass production, standing crop, and composition, the latter either as perennials or Decreasers. Secondary determinants, in varying degrees of importance, are the thickness and base status of the A horizon, distance to permanent drinking water, and competition by woody plants. Herbivore utilization is insignificant or at most, plays a relatively minor role. Herbivores appear to exert a negative influence on Decreaser abundance only when soil moisture stress exceeds a threshold level. When this is exceeded, relatively low herbivore densities are apparently sufficient to reduce Decreaser abundance. The definitions of Decreasers and Increasers consequently require revision to take into account the overriding influence of environmental factors, particularly those of soil moisture stress. The calibration of the disc pasture meter was re-evaluated. The relation between mean disc height and standing crop is non-linear. Up to a mean disc pasture meter height of 260 mm, the correlation between this parameter and above-ground standing crop is very strong (r2 = 0.95; P<0.0005). Beyond this height, the correlation is very poor (r2 =0.09; P<0.0005), apparently being strongly influenced by the structure of the grass plant, with tall grasses, or grasses with highly lignified culms resulting in a weaker correlation.
Grasslands of Umtamvuna Nature Reserve, KwaZulu-Natal : a description and recommendations for monitoring.
(1995) Le Roux, Noel Peter.; Zacharias, Peter John Kenneth.
The main aim of this study was to classify and map the threatened coastal grassland communities of the 3 257 ha Umtamvuna Nature Reserve (UNR) in KwaZulu-Natal (30°07'30" to 300 11'05"E; 30°55'00" to 31°04'30"S). Secondary aims were to relate past management and selected environmental variables to community composition and to develop guidelines for monitoring. Alpha diversity was measured using a Whittaker plot and revealed 119 species. A pilot study to test the efficiency of botanical techniques showed that a point based technique (nearest plant method in a 20 X 20 m plot) was efficient (52 minutes for recording 200 points), but recorded only 23% of the species. By increasing the number of points to six hundred, 34% of the species were recorded in 178 minutes; the same time was required to randomly place 30 quadrats (50 X 50 cm), which revealed 80% of the species. Tests for replicate similarity showed a high retrieval of internal association (PS = 86%), using abundant species only and 100 points per plot. The point based technique was thus efficient in detecting abundant species and was acceptable for producing a classification, especially in this case where a comprehensive species list already existed. Indirect gradient analysis (TWINSPAN) identified six grassland communities. An ordination using detrended correspondence analysis (DCA) contributed towards the community classification and grazing and fire frequency gradients were inferred from this ordination. Protea roupelliae communities were common but did not influence grass species composition . Canonical ordination revealed that, of the eight environmental variables measured, 'distance from the sea' strongly affected species distribution (r= 0.83). Cost effectiveness was considered in the development of a monitoring programme. Point based monitoring techniques favoured by sourveld researchers in KwaZulu-Natal were found to be inefficient, particularly for studies requiring the measurement of both species richness and community composition. Randomly located 100 X 100 cm quadrats, located in selected sites which represent previously identified communities, was more efficient. This study contributed towards a refinement of information on the grassland communities of KwaZulu-Natal and supported the use of point data for the classification of grasslands not previously studied. It also demonstrated that point based techniques were not suitable for meeting all grassland monitoring requirements.
Monitoring serial changes in coastal grasslands invaded by Chromolaena odorata (L.) R.M. King and Robinson.
(2000) Goodall, Jeremy Marshall.; Zacharias, Peter John Kenneth.
The objective of this study was to describe the impacts of the density of Chromolaena odorata (chromolaena) on species composition in coastal grasslands and to investigate serial changes in the vegetation following the implementation of a burning programme. The thesis deals with key ecological concepts and issues, so a comprehensive literature review is included. Chromolaena invades coastal grasslands that are not burnt regularly (i.e. biennially). Grasslands that were not burnt for 30 years were seral to secondary forest. The successional pathway from open grassland to closed canopy forest varied according to soil type. Coastal grasslands on Glenrosa soils were characterised by savanna at an intermediate stage between the grassland and forest states. Shading ended the persistence of savanna species (e.g. Combretum molle, Dichrostachys cinerea and Heteropyxis natalensis) in forest, whereas forest precursors (e.g. Canthium inerme, Maytenus undata and Protorhus longifolia) only established where fire was absent. Chromolaena infestations were characterised by multi-stemmed adult plants of variable height (i.e. 1-3 m), depending on soil type. Regic sands did not support stratified woody vegetation and chromolaena infestations were self-supporting, reaching a maximum height of 1.5 m. Glenrosa soils supported tree communities and chromolaena reached more than 3 m in places. The density of chromolaena affected species composition in grasslands with moderate to dense stands (> 5 adult plants m ¯² or >50000 shrubs ha ¯¹). Chromolaena stands became monospecific when the number of adult plants exceeded 7 m ¯². Succession to forest also ceased once chromolaena became thicket-forming. Fire-induced mortality of the chromolaena depended on grass fuel loads. Grass cover of 30% (c. 1 000 kg ha ¯¹) was required to achieve 80% mortality of the parent infestation after the initial burn. Dense infestations could only be killed by running head-fires from adjacent grasslands into thickets. Under conditions where head-fires could not be used, infestations were slashed and burnt at the height of the dry season (July to August) to achieve an 80% kill rate. Seedlings were killed (99%) by annual burning in sparse (≤ 10000 shrubs ha ¯¹) to moderate < 50 000 shrubs ha ¯¹) infestations. The suppression of chromolaena and other alien species, establishing on bare ground after clearing dense infestations, required chemical control until grass cover was sufficient (i.e. 1 000 kg ha ¯¹) to effect uniform burning. Certain secondary alien invaders (e.g. Lantana camara, Psidium guajava and Solanum mauritianum) persisted by coppicing profusely after fire and herbicides need to be integrated into burning programmes when these species occur. Grasslands on regic sands (e.g. Ischaemum fasciculatum, Panicum dregeanum and Themeda triandra) were more resilient to the modifying effects of woody vegetation, than grasslands on Glenrosa soils. Grasslands on Glenrosa soils did not revert to an open state but persisted as ruderal savanna grassland (e.g. Eragrostis curvula, Hyparrhenia tamba and Cymbopogon validus) once fire-resistant tree species (e.g. Combretum molle and Heteropyxis natalensis) had established. Depending the objectives for land management and the vegetation's condition, coastal grasslands can be rehabilitated and managed in multiple states, i.e. grassland, savanna or forest communities. A state-and-transition model based on the empirical data recorded in the study is presented and shows chromolaena altering vegetation states from open grassland to chromolaena dominated thicket. The model illustrates chromolaena thickets as the dominant phase of a moist coastal forest/savanna succession, irrespective of soil type, in absence of appropriate land management practices (e.g. control burning and integrated control of alien vegetation). This model should aid in planning strategies for the control of chromolaena in subtropical grasslands in South Africa.
Patch grazing at Kroomie.
(2003) Du Toit, Justin Christopher Okes.; Zacharias, Peter John Kenneth.; Danckwerts, Jock Eric.
The patch structure of the grass sward at Kroomie (26°25'38"E 33°48'30"S) in a semi-arid savanna in the Eastern Cape, South Africa, was investigated. The study was conducted on long term grazing trials on five treatments varying in stocking rate (SR; recommended (low) and 1.5 x recommended (high), grazing system (continuous and rotational), and animal type (cattle and sheep). The treatments studied were CR (cattle, rotational stocking, low SR), CC (cattle, continuous stocking, low SR), CH (cattle, rotational stocking, high SR), SC (sheep, continuous stocking, low SR), and SR (sheep, rotational stocking, low SR). Rainfall during the two years of the study (1997/98 and 1998/99) was slightly below the mean average rainfall of the area (66 and 84% of the mean of 519 mm, respectively). Analysis of sward height data using Maximum Likelihood Estimation reflected a bimodal height structure in all treatments. Due to a high overlap of the two distributions in some cases, however, the height at which to separate patches (short grass) from non-patches (tall grass) could not be determined. Canonical Correspondence Analysis (CCA) was used to relate species composition to sward height. It emerged that there are two distinct grass communities at Kroomie, and these are associated with sward height (i.e. patches and non-patches). The interface (in cm) between these two communities, as determined using Two Way Indicator Species Analysis (TWINS PAN) was 6 cm, and this value was subsequently used to discriminate between patches and non-patches. Sward structure was affected by treatments. Animals (cattle and sheep) stocked rotationally at low SR's grazed less than a third of the total area, and this grazing was concentrated primarily in small patches (< 6 m; length is used as a linear indicator of patch size). Animals stocked continuously at low SR's grazed approximately half the area, in small and large (up to 40 m) patches. Animals in the CH treatment grazed approximately two-thirds of the area, in both small and large patches. There was an inverse relation between the size of patches and the size of non-patches, as expected. Nine common grass species were related to sward height. Digiteria eriantha, Eragrostis racemosa, Eustachys paspaloides, and Microchloa caffra were associated with short swards, while Cymbopogon pluronodis, Eragrostis chloromelas, and Sporobolus fimbriatus were associated with tall swards. Themeda triandra (themeda), the most abundant grass at Kroomie, was principally associated with tall swards, but was present at all sward heights. Applying CCA demonstrated a considerable difference between the species composition of patches and of non-patches. There was also a difference in composition between treatments, although these were not as pronounced. Patches reflected a higher species diversity than non-patches. There was a significant (P<0.05) effect of treatment, and of an interaction of treatment by sward structure (i.e. patches and non-patches), on the density of themeda plants. The density of themeda plants was positively correlated with patch size, which suggested that themeda plants that have been grazed may suffer fatal competition from ungrazed neighbours. Anecdotal evidence suggested that patches are stable over the medium term, and that non-patches that are grazed during a drought return to a non-patch structure after rainfall. There was no evidence to support the contention that rotational stocking reduced patch-selective grazing, nor that the species composition of rotationally stocked treatments was better than continuously stocked treatments.
Re-vegetation dynamics of land cleared of Acacia mearnsii (black wattle)
(2005) Glaum, Melanie Jane.; Zacharias, Peter John Kenneth.; Granger, James Edmund.; Everson, Theresa Mary.; Smith, M.
The overall aim of the study was to investigate re-vegetation of disturbed sites, using nursery grown plugs (from seedling trays) of Themeda triandra, Heteropogon contortus and Hyparrhenia dregeana in order to reach practical management guidelines for re-vegetation using indigenous grass plugs. A number of field trials were set up at Kamberg Nature Reserve (29°24'S, 29°40'E) on a site that was clear felled of A. mearnsii in October 1997. The trials were established in January 1998 and January 1999. A total of approximately 52 ,000 nursery raised plugs of T. triandra, H. contortus and H. dregeana were planted into an area of approximately 7,000 m2 . In the planting density trial , plugs of H. dregeana only and a combination of T. triandra/H. contortus were planted at 15 cm and 30 cm spacings. The T. triandra/H. contortus combination at 30 spacing showed the greatest survival and lateral plant growth (tiller number and basal area) and this combination is thus recommended. In the over-sowing trials, the H. dregeana and T. triandra/H. contortus combination at both 15 cm and 30 cm spacing were over-sown with E. curvula. The survival and lateral growth of the T. triandra/ H. contortus combination at 30 cm was again greater than the other treatments. Over-sowing with E. curvula suppressed the survival and lateral growth of the planted plugs across all treatments compared to not over-sowing. The over-sown conditions showed a significant decrease in the diversity of the plots, both in the number of species present and the Shannon diversity index. An area that had been cleared of A. mearnsii and sown to E. curvula 25 years previously was shown to have a lower number of species than the neighbouring veld. Nursery raised plugs of T. triandra were planted into the mature E. curvula in an attempt to improve the biodiversity of these areas. To re-introduce T. triandra into these E. curvula swards the plugs must be planted into the centre of a gap rather than around the base of an E. curvula plant. For improved survival of the plugs the E. curvula tufts must be clipped, while for best lateral growth the E. curvula tufts must be sprayed with a glyphosate herbicide three months prior to planting and clipping. However, the added expense of spraying and clipping is not warranted as the clipped treatments also showed good growth. Transplant shock is common when planting nursery raised plugs out into the field, as there is a relatively small root volume in the plug compared to the above ground leaf biomass. Alleviation of moisture stress at planting using a starch based polymer with high water holding capacity (Terrasorb®) and a white, needle punched geo-fabric (Agrilen®) to provide a seven day period of artificial shade after planting did not show significant improvements over the control with regards to survival or plant growth. Thus these methods of moisture amelioration are not recommended in revegetation through planting of plugs at this study site. A trial was established to investigate the biomass production of six different treatments to determine their potential to support a fire. The total biomass for the plots which were over-sown by E. tef and planted to only H. dregeana were on average sufficient for a fire, but there was a discontinuous fuel load across these plots, especially in the replications that had very low survival rates and thus these plots could not be burnt. The control and herbicide sprayed plots also showed sufficient fuel load for a fire, but this fuel load was made up of A. mearnsii saplings and bramble with very little grass cover and thus a fire would not have burnt through these plots either. The T. triandra/H. contortus combination did not produce sufficient fuel load, due to poor survival. Thus only the plots over-sown with E. curvula were able to burn in this trial and as a burning trial per se the trial was abandoned. Seed bearing hay (thatch) was collected in early summer (December 1997) and late summer (April 1998). Both times of year of harvesting proved to be successful in terms of grass cover, although the early harvested thatch had a greater number of species per plot. The Shannon diversity indexes of the two treatments were not significantly different. The multi-response permutation procedure technique confirmed that there was a compositional difference between the treatments. By the end of the trial Harpochloa falx and T. triandra and H. dregeana were indicators for the early and the late harvested thatch respectively. Comparing the thatching trial and the planting density trial indicated that the T. triandra/H. contortus combination at 30 cm spacing would be recommended to maximize biodiversity. The summer months have been shown to be the best time to plant the plugs, although the actual success will be dependant on the conditions within a particular year. The plugs should not be kept in the nursery for longer than three months and larger plugs (96 seedlings per tray) should be used. Nursery raised plugs of T. triandra and H. contortus were planted in an equal mix in an area that was cleared of A. mearnsii in 1996. By June 1998 661 H. contortus seedlings and 14 T. triandra seedlings had germinated naturally. The November 1998 population consisted of 418 H. contortus seedlings and 18 T. triandra seedlings. By May 2000 the June 1998 population showed a survival of 78.4% and the November 1998 population showed a survival of 91 .1 %. In the various trials, the ability of the nursery raised plugs used for re-vegetation to suppress the regrowth of A. mearnsii was investigated by determining the number of A. mearnsii seedlings per metre squared. The plant spacing and species of plugs used did not have a significant effect on the number of A. mearnsii seedlings per metre squared. Over-sowing with E. curvula did, however, significantly suppress the wattle re-growth. In the thatching trial the early harvested plots showed lower numbers of A. mearnsii per metre squared than the late harvest plots, as they were covered with a thick layer of thatch soon after the A. mearnsii was cleared which suppressed the A. mearnsii re-growth. Although E. curvula is able to produce a high biomass and suppress the A. meansii seedlings, it has a detrimental effect on the biodiversity of the area. Therefore, in conservation areas, where biodiversity is of great importance the planted plugs (at 30 cm spacing) or seed bearing hay must be used in preference to sowing E. curvula , although it must be remembered that greater follow up control is likely to be needed with planted plugs or seed bearing hay. The area must be planted or thatched as soon as possible after clear felling to provide competition for the A. mearnsii seedlings.
The seasonal patterns in plant quality in various ecological zones in Natal.
(1990) Zacharias, Peter John Kenneth.; Tainton, Neil M.
The objectives of this study required that the following investigations be undertaken: 1) to determine if plant quality can be altered by modifying growing conditions; 2) to quantify the seasonal trends in plant quality from different sites; 3) to relate differences between sites to environmental variables; 4) to develop an objective classification of SWEETNESS; and 5) to plan future research. The majority of the commercial and subsistence livestock in southern Africa rely almost entirely on veld (rangeland) for their supply of nutrients. These rangelands are traditionally and conventionally managed according to their classification as 'SWEETVELD' and 'SOURVELD'. An intermediate group 'MIXED-VELD' is also recognised. The subjective classification is based on the quality (nutritive value) of the rangeland when it is mature (winter). Both extremes of 'sweet' and 'sour' rangeland contain many of the same species and this thesis considers the relationship between the soil ( chemical and physical) and the physical environment and plant quality of a single indicator species Themeda triandra (red grass). A glass house experiment was used to determine the effect of manipulating the soil environment on the quality of T. triandra. There were no significant differences ( P> 0. 05) between any of the six treatments (combinations of eutrophic, ameliorated and dystrophic soil together with 'sweet' and 'sour' T. triandra plan ts) . When compared as a group significantly (PsO. 01) higher in disappearance ( CDMD) than the SOUR group. the SWEET group were cellulase dry matter However the difference was only 1 . 5% CDMD uni ts and is believed to be biologically unimportant. It was concluded that the quality of T. triandra can not be altered by manipulating its growing conditions. In a field investigation the seasonal pattern and the relationship between environmental variables and plant CDMD, N, P, K, Mg, Ca, Sand Zn are described for Natal, South Africa. Most models have significant (Ps0.01) R2 values but very few show any strong relationship between soil chemical status and forage j~-l quality. ALTITUDE appears most in the models as a related variable. The models would have little predictive value outside Natal and do not contribute or describe adequately the factors determining seasonal patterns in plant quality at different locations. A multivariate approach is used to provide an objective index of 'SWEETNESS' (based on seasonal variations in plant quality at 31 sites over 23 months), and this is related to environmental variables. This analysis also showed that the soil environment was only weakly related to plant quality. The results are confusing given the wide variations in both plant quality and soil chemistry in the data presented.
A strategy for optimal beef production off sourveld.
(2003) Buntting, Clive Bartle.; Zacharias, Peter John Kenneth.
The economic necessity of a better production strategy on sourveld promoted this study. Production of marketable two-tooth steers in the summer season and overwintering of all cattle without excessive feed costs were motivating factors. The problems of economic beef production were identified as resulting from the seasonal flow of forage quality from sourveld grass production. Season long rests, early burning and non-selective grazing of nutritious grass were identified as essential elements of a new utilization strategy. A 'forage reserve', built into the system to cater for fluctuations in grass production due to varied climatic conditions, is used as an indicator of the seasonal stocking rate. This provides a barometer in relation to the economic and ecological carrying capacity of the property. Research was conducted on the winter utilization of rested veld and its effects on grass species composition and vigour in the following season. It was found that the winter grazing of the rested veld did not affect (P>0.05) the subsequent production in the three seasons of this study on 'Stratherne' in the Dundee district, KwaZulu-Natal South Africa (30°17'E 28°17'S). The grass species composition of four transects was recorded in 1994, prior to the implementation of the grazing system under test. The same transects were recorded again in 2002 to determine the effect of the change in utilization on grass species composition. It was found that a more productive state was developing in response to the strategy implemented in this study. The general trend has been for sites to move from a Hyparrhenia hirta dominated state to a more productive one associated with species such as Themeda triandra. Summer mass gains of steers (147 kg and 143 kg over the two summers) have improved over the previous systems applied (average 119 kg), as a result of the more nutritious grazing. A greater proportion reached market readiness as two-year old to two and half year olds (97%) on veld, which is far superior to the 38% quoted from research using similar Bonsmara type steers from 'conventional' systems. Monitoring and flexibility are important in the application of the strategy to conditions in Africa. The principles of adaptive management (monitoring, recording, constant learning and adaptation) will build a data base to ensure long-term success of the strategy. A change of focus in grazing strategy from needs of animals to the needs of plants is strongly advocated.
Summer supplementation of beef cattle on veld and kikuyu pastures.
(2004) Reynolds, Richard Norman.; Darroch, Mark Andrew Gower.; Zacharias, Peter John Kenneth.
In KwaZulu-Natal the production of beef in summer from veld is a common enterprise. Many techniques are available to optimise the productivity of this enterprise, from improving the quality of the grazing resource (planted pastures) to improving the diet of animals using nutritional supplements. To gain an insight into the production potential and financial returns possible from such improvements a trial was established at the Ukulinga Research Farm during two consecutive summer growing seasons (1997-1998 and 1998-1999). The aim was to determine the optimum beef production system for the area from both veld and kikuyu pastures. To evaluate the benefits of supplying supplementary nutrition, four alternate feed supplements, namely: 1) a Standard commercially available molasses-based protein/mineral/energy supplement; 2) a Brewers grain based protein/mineral/energy supplement, and two supplements consisting of the Brewers grain ration with either; 3) Avoparcin (an additive that improves dietary energy) or 4) Bentonite (an additive that increases the bypass of protein) were compared. As stocking rate has been shown to influence the quality of the diet consumed, the kikuyu pasture was grazed at both the recommended (1.92 LSD ha¯¹) and half the recommended stocking rate for the region. In addition, a commercial hormonal implant was applied to half of the cattle in each treatment. Grazing was monitored using the falling plate disc meter to measure pasture bulk density and laboratory analyses of herbage grab samples for digestibility and crude protein percentage. Cattle were weighed on a weekly basis and their condition was scored prior to slaughter. All enterprise costs and returns were recorded to facilitate financial analyses of the five treatments. Low rainfall and high midsummer temperatures had a detrimental effect on the productivity of the grazing and hence it was difficult to optimise production in either season. In the first season, a midsummer drought decreased the quality and quantity of both veld and kikuyu, limiting mass gain during the latter part of the season. A delay in the onset of rain at the start of the second season limited the available grazing season to 121 days as opposed to 154 days, though fodder production during the season was not limited. Trends in herbage production (quality and quantity) from veld showed moderate quality (Crude protein 7.02%; digestibility 50.2%) with an average available herbage of 1670 kg DM ha¯¹. As anticipated, kikuyu had higher quality (Crude protein 10.84%; digestibility 53.5%) and available herbage (2730 kg DM ha¯¹). These results were similar to regional benchmarks. The variable rainfall highlighted both the drought tolerance of veld and the minimum water requirements of kikuyu pastures. Lighter stocking rates tended to reduce the negative impact of moisture stress on Kikuyu pastures. The best method of producing beef (averaged over two seasons) was from heavily stocked Kikuyu pastures using the Standard supplement (1107.63 kg livemass ha¯¹). Cattle grazing veld and utilising the Avoparcin supplement produced beef at a rate of 95.96 kg ha¯¹. In comparison, the unsupplemented cattle grazing Kikuyu produced 834.87 kg ha¯¹, whilst veld grazing produced 64.43 kg ha¯¹. Hormonal implants significantly (P≤0.05) improved beef production from all sources of grazing. A lack of rain limited grazing time, causing all the cattle to be marketed whilst too lean - this negatively affected live mass gain and, hence, net financial. Although improved biological production is desirable, it is important to ensure that these gains are financially sustainable. Within the trial environment, implanted cattle fed the Standard supplement and grazing Kikuyu pastures at a high stocking rate provided the highest average gross margin of R 859.59 ha¯¹. Changing to this production system from unsupplemented veld improved expected profit by R 632.58 ha¯¹ (averaged over both seasons). Further financial analyses indicated that beef purchase price had the greatest influence on the added profit from switching from the control treatment. From a scientific standpoint these data are conclusive but it is important to remember that consumer pressure and concerns can often limit the introduction of production improvements. Such is the case with both hormonal implants and antibiotic feed additives (Avoparcin) although considering the impact of such limitations is speculative and beyond the scope of this trial.
Towards a predictive understanding of savanna vegetation dynamics in the eastern Lowveld of South Africa : with implications for effective management.
(2005) Peel, Michael John Stephen.; Zacharias, Peter John Kenneth.; Biggs, Chara.
The purpose of this study was to develop and test a predictive understanding of the vegetation dynamics of the Lowveld of South Africa (30°35'E to 30°40'E and 24°00'S to 25°00'S). The study covered about 5000 km2 in Adjacent Private Protected Areas (APPA) adjoining the Kruger National Park (KNP). Data gathering (800 sites; 23 properties) commenced in 1989 and those recorded up to 2004 are reported here. The value, both ecological and economic, of the wildlife and tourism industry dependent on this savanna region is discussed in both historical and current perspectives. A range of land-use objectives and anthropogenic interventions were exposed. The properties ranged in size from 30 to 800 km2 and formed an effective and extensive manipulative experiment for investigating interaction of bush density, animal stocking, use of fire and landscape-scale processes. The first descriptive classification (at 1:250 000) of the area was developed using Inverse Distance Weighted interpolations. This confirms similar landscape/vegetation patterns in the KNP and Mocambique. The current mode of determining stocking density or carrying capacity was interrogated and indices suitable for complex multi-species systems developed. This was done in the context of equilibrial/disequilibrial paradigms. Application of the original indices resulted in drought-related declines in animal biomass of 4000 kg km2 over 20 years due to overestimation of carrying capacity. The model proposed here uses rainfall, animal type, biomass and vegetation parameters to determine stocking density for both coarse (regional) and ranch-specific scales. Principal driving determinants (rainfall, geology, soil type, tree density canopy cover, animal numbers, feeding classes and fire) of vegetation structure and their influence on the herbaceous layer were investigated. Groupings on ecological potential showed 'high' potential areas are less sensitive to animal impact than those classified as 'low' potential. Sustainability, embedded in a forward-looking component viz. Strategic Adaptive Management (SAM) with well-articulated endpoints viz. Thresholds of Potential Concern (TPCs) was used to study fluctuations in animal populations with Connochaetes taurinus (Blue wildebeest) as the case study. The TPC approach provides strong pointers for proactive management aimed at maintaining the system within bands defined by TPCs supporting operationally practical and periodically reviewed objectives.